The pairs of high and low horns tend to act as two duets. This can be seen in some music like Brahms were Horns 1 and 2 are crooked in Bb, and horns 3 and 4 are crooked in F. This way the horn section can form more chords and play a wider variety of melodies in different keys.
Now I bet I'm still missing something interesting on this topic and if you'd like to contribute and voice your opinion either leave a comment with your thoughts, or you can shoot me an email at austinbaugh@gmail.com.
Thanks!

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